一、微生物的生長和繁殖
weishengwuzaishiyidehuanjingtiaojianxia,buduandixishouyingyangwuzhi,binganzhaozijidedaixiefangshijinxingdaixiehuodong,ruguotonghuazuoyongdayuyihuazuoyong,zexibaozhideliangbuduanzengjia,tijideyijiada,yushibiaoxianweishengchang。jiandandishuo,shengchangjiushiyoujitidexibaozufenyujiegouzailiangfangmiandezengjia。
danxibaoweishengwuruxijun,shengchangwangwangbansuizhexibaoshumudezengjia。dangxibaozengchangdaoyidingchengdushi,jiuyierfenliefangshi,xingchenglianggejibenxiangsidezixibao,zixibaoyouzhongfuyishangguocheng。zaidanxibaoweishengwuzhong,youyuxibaofenlieeryinqidegetishumudezengjia,chengweifanzhi。zaiyibanqingkuangxia,danghuanjingtiaojianshihe,shengchangyufanzhishizhongshijiaotijinxingde。congshengchangdaofanzhishiyigeyouliangbiandaozhibiandeguocheng,zhegeguochengjiushifayu。(附圖是大腸杆菌 E.coli.的分裂繁殖,可是費了俺好大的勁。^-^)
微生物處於一定的物理、化學條件下,生長發育正常,繁殖速率也高;如果某一或某些環境條件發生改變,並超出了生物可以適應的範圍時,就會對機體產生抑製乃至殺滅作用。
二、細菌純培養的群體生長規律
大多數細菌的繁殖速度都很快,大腸杆菌在適宜條件下,每20分鍾左右便可分裂一次,如果始終保持這樣的繁殖速度,一個細菌在48小時內,其子代群體將達到無法想象的數量。然而,實際情況並非如此。

將少量單細胞純培養接種到一恒定容積的新鮮液體培養基中,在適宜的條件下培養,定時取樣測定其細菌含量,可以看到以下現象:kaishiyouyiduanzanshijian,xijunshuliangbingbuzengjia,suizhixijunshumuzengjiahenkuai,jierxijunshuyouquwending,zuihouzhujianxiajiang。ruguoyipeiyangshijianweihengzuobiao,yixijunshumudeduishuhuoshengchangsuduweizongzuobiaozuotu,keyidedaoyitiaoquxian,chengweifanzhiquxian,tongchangyouchengweishengchangquxian。shengchangquxiandaibiaolexijunzaixindeshiyidehuanjingzhongshengchangfanzhizhizhishuailaosiwangquanguochengdedongtaibianhua。genjuxijunshengchangfanzhisulvdebutong,kejiangshengchangquxiandazhifenweiyanchiqi、對數期、穩定期和衰亡期四個階段。
(1)延遲期:shaoliangxijunjiezhongdaoxinxianpeiyangjihou,yibanbulijijinxingfanzhi,shengchangsudujinyuling。yincizaikaishiyiduanshijian,xijunshujihubaochibubian,shenzhishaoyoujianshao。zheduanshijianbeichengweiyanchiqi,youchengweichihuanqi、調整期或滯留適應期。處於延遲期細菌細胞的特點是分裂遲緩、代謝活躍。延遲期的長短與菌種、種齡、接種量和培養基成分有關。
(2)對數期:對(dui)數(shu)期(qi)又(you)稱(cheng)指(zhi)數(shu)期(qi)。這(zhe)一(yi)階(jie)段(duan)突(tu)出(chu)特(te)點(dian)是(shi)細(xi)菌(jun)數(shu)以(yi)幾(ji)何(he)級(ji)數(shu)增(zeng)加(jia),代(dai)時(shi)穩(wen)定(ding),細(xi)菌(jun)數(shu)目(mu)的(de)增(zeng)加(jia)與(yu)原(yuan)生(sheng)質(zhi)總(zong)量(liang)的(de)增(zeng)加(jia),與(yu)菌(jun)液(ye)混(hun)濁(zhuo)度(du)的(de)增(zeng)加(jia)均(jun)呈(cheng)正(zheng)相(xiang)關(guan)性(xing)。
(3)穩定期:又you稱cheng恒heng定ding期qi或huo最zui高gao生sheng長chang期qi。處chu於yu穩wen定ding期qi的de微wei生sheng物wu,新xin增zeng殖zhi的de細xi胞bao數shu與yu老lao細xi胞bao的de死si亡wang數shu幾ji乎hu相xiang等deng,整zheng個ge培pei養yang物wu中zhong二er者zhe處chu於yu動dong態tai平ping衡heng,此ci時shi生sheng長chang速su度du又you逐zhu漸jian趨qu向xiang零ling。
穩定期的細胞內開始積累貯藏物,如肝糖、異染顆粒、脂肪粒等,大多數芽孢細菌也在此階段形成芽孢。如果為了獲得大量菌體,就應在此階段收獲,因這時細胞總數最高;這一時期也是發酵過程積累代謝產物的重要階段,某些放線菌抗生素的大量形成也在此時期。
(4)衰亡期:穩定期後如再繼續培養,細菌死亡率逐漸增加,以致死亡數大大超過新生數,群體中活菌數目急劇下降,出現了“負生長”,此階段叫衰亡期。
圖2:細菌生長的典型曲線
(Ⅰ.延遲期, Ⅱ.對數期, Ⅲ.穩定期, Ⅳ.衰亡期)
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